Pituophis

Pituophis
Pacific gopher snake Pituophis catenifer catenifer
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Squamata
Suborder:	Serpentes
Family:	Colubridae
Tribe:	Lampropeltini
Genus:	Pituophis Holbrook, 1842
Synonyms
Churchilla, Elaphis, Epiglottophis, Pityophis, Rhinechis

Pituophis is a genus of nonvenomous colubrid snakes, commonly referred to as gopher snakes, pine snakes, and bullsnakes, which are endemic to North America.

Nomenclature

The genus name Pituophis is a Latinized modern scientific Greek compound Πιτυόφις : "pine snake"; from Ancient Greek: πίτυς (pítus, "pine"), and Ancient Greek: ὄφις (óphis, "snake").

Geographic range

Species and subspecies within the genus Pituophis are found throughout Mexico, the Southern and Western United States and Western Canada.

Description

All species of Pituophis are large and powerfully built. The head is relatively small in proportion to the body and it is only slightly distinct from the neck. The rostral is enlarged and elongated, imparting a characteristic somewhat pointed shape to the head. All the species occurring in the United States have four prefrontals instead of the usual two.

Modified epiglottis

In all snakes of the genus Pituophis, the epiglottis is peculiarly modified so that it is thin, erect and flexible. When a stream of air is forced from the trachea, the epiglottis vibrates, thereby producing the peculiarly loud, hoarse hissing for which bullsnakes, gopher snakes and pine snakes are well known.

Species and subspecies

Image	Scientific name	Common Name	Subspecies	Distribution
	Pituophis catenifer (Blainville, 1835)	gopher snake	P. c. affinis (Hallowell, 1852) – Sonoran gopher snake P. c. annectens Baird & Girard, 1853 – San Diego gopher snake P. c. bimaris Klauber, 1946 – central Baja California gopher snake P. c. catenifer (Blainville, 1835) – Pacific gopher snake P. c. coronalis Klauber, 1946 – Coronado Island gopher snake P. c. deserticola Stejneger, 1893 – Great Basin gopher snake P. c. fulginatus Klauber, 1946 – San Martin Island gopher snake P. c. pumilis Klauber, 1946 – Santa Cruz Island gopher snake P. c. sayi (Schlegel, 1837) – bullsnake	North America
	Pituophis deppei (A.M.C. Duméril, 1853)	Mexican bullsnake	P. d. deppei (A.M.C. Duméril, 1853) – southern Mexican pine snake P. d. jani (Cope, 1861) – northern Mexican pine snake	central, Mexico
	Pituophis insularis Klauber, 1946	Credos Island gopher snake		Isla de Credos, Mexico
	Pituophis lineaticollis (Cope, 1861)	Middle American gopher snake	P. l. gibsoni Stuart, 1954 P. l. lineaticollis (Cope, 1861)	From Mexico city, south through Mexico and to Guatemala
	Pituophis melanoleucus (Daudin, 1803)	pine snake	P. m. lodingi Blanchard, 1924 – black pine snake P. m. melanoleucus (Daudin, 1803) – northern pine snake P. m. mugitus Barbour, 1921 – Florida pine snake	southeastern United States
	Pituophis ruthveni Stull, 1929	Louisiana pine snake		west-central Louisiana and East Texas
	Pituophis vertebralis (Blainville, 1835)	Cape gopher snake		southern Baja California Sur, Mexico.

Further reading

• Holbrook, John Edwards (1842). North American herpetology : or, A description of the reptiles inhabiting the United States. Vol. IV. Philadelphia: J. Dobson. p. 7.

External links

• Genus Pituophis at The Reptile Database
• Accounts of four subspecies, P. deppei deppei, P. deppei jani, P. lineaticollis lineaticollis and P. lineaticollis gibsoni, are given in Duellman WE (1960). "A Taxonomic Study of the Middle American Snake, Pituophis deppei". University of Kansas Publications, Museum of Natural History 10: 599–610.